IOC World Bird List 2.4

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ORDER

FAMILY

NOTES ON CLASSIFICATION

Tinamiformes

Tinamous

Tinamidae

"Early Bird" DNA analyses suggest that flighted tinamous may have evolved from flightless ratites (Hackett et al. 2008)

Struthioniformes

Ostriches

Struthionidae

 

Rheiformes

Rheas

Rheidae

 

Casuariiformes

Cassowaries

Casuariidae

 
 

Emu

Dromaiidae

 

Apterygiformes

Kiwis

Apterygidae

 

Galliformes

Megapodes

Megapodiidae

 
 

Chachalacas, Curassows & Guans

Cracidae

 
 

Guineafowl

Numididae

 
 

New World Quail

Odontophoridae

 
 

Pheasants, Fowl & Allies

Phasianidae

 

Anseriformes

Screamers

Anhimidae

 
 

Magpie Goose

Anseranatidae

 
 

Ducks, Geese & Swans

Anatidae

 

Sphenisciformes

Penguins

Spheniscidae

 

Gaviiformes

Loons

Gaviidae

 

Procellariiformes

Albatrosses

Diomedeidae

 
 

Petrels, Shearwaters

Procellariidae

 
 

Storm Petrels

Hydrobatidae

The subfamilies Hydrobatinae and Oceanitinae may not be sister taxa and therefore be separate families (Nunn and Stanley 1998; Hackett et al. 2008; Christidis and Boles 2008)

 

Diving Petrels

Pelecanoididae

 

Podicipediformes

GrebesGGGrebes

Podicipedidae

 

Phoenicopteriformes

Flamingos

Phoenicopteridae

 

Phaethontiformes

Tropicbirds

Phaethontidae

The tropicbirds (Phaethontidae) do not belong in the Pelecaniformes and merit treatment as the Order Phaethontiformes (Hackett et al. 2008, Christidis and Boles 2008)

Ciconiiformes

Storks

Ciconiidae

The Ciconiiformes now includes only one family; the storks (Ciconiidae); ibises and spoonbills (Threskiornithidae) and herons (Ardeidae) are are moved the Pelecaniformes (Hackett et al 2008)

Pelecaniformes

Ibises, Spoonbills

Threskiornithidae

Ibises and spoonbills (Threskiornithidae) and herons (Ardeidae) are sister taxa in the Order Pelecaniformes (Hackett et al. 2008)

 

Herons, Bitterns

Ardeidae

 
       
 

Hamerkop

Scopidae

 
 

Shoebill

Balaenicipitidae

 
 

Pelicans

Pelecanidae

 
Suliformes

Frigatebirds

Fregatidae

 
 

Gannets, Boobies

Sulidae

 
 

Cormorants, shags

Phalacrocoracidae

 
 

Anhingas, darters

Anhingidae

 

Accipitriformes

New World Vultures

Cathartidae

SACC elevates the New World Vultures (Cathartidae) to a separate order, the Cathartiformes

 

Secretarybird

Sagittariidae

Secretarybird is best treated as a monotypic family Sagittariidae in accord with the Tree of Life phylogeny (Hackett et al. 2008)

 

Ospreys

Pandionidae

Ospreys are separated to the family Pandionidae in accord with the Tree of Life phylogeny (Hackett et al. 2008)

 

Kites, Hawks & Eagles

Accipitridae

 

Falconiformes

Caracaras, Falcons

Falconidae

The falcons (Falconidae) are not members of the Accipitriformes, but are a separate lineage and Order more closely related to parrots and in turn the Passeriformes (Hackett et al. 2008). Resequence in the future.

Otidiformes

Bustards

Otididae

Move bustards (Otididae) from the Gruiformes to their own Order, the Otidiformes (Hackett et al. 2008)

Mesitornithiformes

Mesites

Mesitornithidae

Move mesites (Mesitornithidae) from the Gruiformes to their own Order, the Mesitornithiformes (Hackett et al. 2008)

Cariamiformes

Seriemas

Cariamidae

Move seriemas (Cariamidae)  from the Gruiformes to their own Order, the Cariamiformes (Hackett et al. 2008)

Eurypygiformes

Kagu

Rhynochetidae

The Kagu (Rhynochetidae) and Sunbittern (Eurypygidae) are sister families removed from the Gruiformes to a new Order, tentatively named the "Eurypygiformes" (Hackett et al. 2008).

 

Sunbittern

Eurypygidae

 

Gruiformes

Flufftails

Sarothruridae

Long recognized to be an artificial assemblage of taxa, the Gruiformes is now restricted to flufftails, finfoots, rails, trumpeters, cranes, and Limpkin (Hackett et al. 2008)

 

Finfoots

Heliornithidae

The finfoots (Heliornithidae) and flufftails (Sarothruridae) are sister taxa (Hackett et al. 2008)

 

Rails, Crakes & Coots

Rallidae

 
 

Trumpeters

Psophiidae

 
 

Cranes

Gruidae

 
 

Limpkin

Aramidae

 

Charadriiformes

Buttonquails

Turnicidae

The buttonquails (Turnicidae) belong in the Charadriiformes, not the Gruiformes (Hackett et al. 2008)

 

Stone-curlews, Thick-knees

Burhinidae

 
 

Sheathbills

Chionidae

 
 

Magellanic Plover

Pluvianellidae

The Magellanic Plover (Pluvianellus) is most closely related to the sheathbills (Chionidae) but treated as a monotypic family by the SACC.

 

Oystercatchers

Haematopodidae

 
 

Crab-plover

Dromadidae

 
 

Ibisbill

Ibidorhynchidae

 
 

Stilts, Avocets

Recurvirostridae

 
 

Plovers

Charadriidae

The golden plovers (Pluvialis) are a separate lineage that may merit family status (Baker et al 2007, TOL)

 

Egyptian Plover

Pluvianidae

The Egyptian Plover (Pluvianus) does not belong to the Glareolidae. It is a separate lineage that is the outgroup to plovers, ibisbill, stilts and their allies (Hackett et al. 2008). Therefore separated here to its own family tentatively named "Pluvianidae"

 

Painted Snipes

Rostratulidae

 
 

Jacanas

Jacanidae

 
 

Plains-wanderer

Pedionomidae

 
 

Seedsnipes

Thinocoridae

 
 

Sandpipers, Snipes

Scolopacidae

 
 

Coursers, Pratincoles

Glareolidae

 
 

Gulls, Terns & Skimmers

Laridae

The skimmers and terns are embedded among the gulls in a monophyletic clade; separation of the families Rynchopidae and Sternidae would render the gull family Laridae paraphyletic relative to these other two groups (Baker et al. 2008).

 

Skuas

Stercorariidae

 
 

Auks

Alcidae

 

Pteroclidiformes

Sandgrouse

Pteroclididae

The sandgrouse (Pteroclididae) are treated as a separate Order Pteroclidiformes because their relationships to other modern birds remain uncertain.

Columbiformes

Pigeons, Doves

Columbidae

 

Psittaciformes

New Zealand Parrots

Strigopidae

Parrots (Psittaciformes) are the sister group to the Passeriformes (Hackett et al. 2008); resequence in future. The New Zealand parrots are a basal lineage that is sister to all other parrots; elevate to family status (Christidis & Boles (2008)

 

Cockatoos

Cacatuidae

Cockatoos are treated as a family separate from other parrots (Psittacidae) following Christidis & Boles (2008)

 

Parrots

Psittacidae

 

Opisthocoformes

Hoatzin

Opisthocomidae

 

Musophagiformes

Turacos

Musophagidae

 

Cuculiformes

Cuckoos

Cuculidae

 

Strigiformes

Barn Owls

Tytonidae

 
 

Owls

Strigidae

 

Caprimulgiformes

Frogmouths

Podargidae

Defer recognition of the Asian frogmouths (Batrachosomus) vs Australasian frogmouths (Podargus) as separate families (Christidis and Boles 2008) until (basal?) relationships of Rigidipenna inexpectata are resolved (see Tree of Life)

 

Oilbird

Steatornithidae

 
 

Potoos

Nyctibiidae

 
 

Nightjars

Caprimulgidae

The eared nightjars (Eurostopodus) are the sister taxon to rest of the Caprimulgidae (Barrowclough et al 2006, Larsen et al. 2007), but they are not equivalent in rank to other families in the Caprimulgiformes.

Apodiformes

Owlet-nightjars

Aegothelidae

The owlet-nightjars (Aegothelidae) are the sister taxon to treeswifts and in turn to the swifts and hummingbirds; hence are included in the Apodiformes (Hackett et al. 2008, Christidis and Boles 2008)

 

Treeswifts

Hemiprocnidae

 
 

Swifts

Apodidae

 
 

Hummingbirds

Trochilidae

 

Coliiformes

Mousebirds

Coliidae

 

Trogoniformes

Trogons

Trogonidae

 

Leptosomatiformes

Cuckoo Roller

Leptosomidae

The Cuckoo Roller is the only member of an ancient lineage (not Coraciiformes) with no known relatives among modern birds and thus put into a separate Order, Leptosomiformes (Hackett et al. 2008)

Coraciiformes

Rollers

Coraciidae

 
 

Ground Rollers

Brachypteraciidae

 
 

Kingfishers

Alcedinidae

The kingfishers (Alcedinidae) comprise three monophyletic clades (Alcedininae, Cerylinae, and Daceloninae) (Moyle 2007); some taxonomists elevate them to family status (Sibley and Monroe 1990, Christidis and Boles 2008)

 

Todies

Todidae

 
 

Motmots

Momotidae

 
 

Bee-eaters

Meropidae

 

Bucerotiformes

Hoopoes

Upupidae

Separation of the hoopoes and hornbills as the Order Bucerotiformes maintains monophyly of the Coraciiformes without submerging the Piciformes and Trogoniformes into the Coraciiformes (Hackett et al. 2008).

 

Wood Hoopoes

Phoeniculidae

The hoopoes (Upupidae) and wood hoopoes (Phoeniculidae) are sister taxa, related in turn to the hornbills (Hackett et al. 2008)

 

Hornbills

Bucerotidae

The hornbills (Bucerotidae) and ground hornbills (Bucorvidae) are sister taxa, related in turn to the hoopoes and wood hoopoes (Hackett et al. 2008)

 

Ground Hornbills

Bucorvidae

 

Piciformes

Jacamars

Galbulidae

The jacamars (Galbulidae) and puffbirds (Bucconidae) are well-established sister taxa (SACC, Hackett et al. 2008), with evidence favoring inclusion in the Piciformes, rather than a separate Order Galbuliformes (TOL, Ericson et al. 2006; cf SACC, Fain & Houde 2004)

 

Puffbirds

Bucconidae

 
  New World Barbets Capitonidae Follow AOU in elevating to Family status as a member of trichotomy with Toucan Barbet and toucans
  Toucan Barbet Semnornithidae Follow AOU in elevating to Family status as a member of trichotomy with New World barbets and toucans
 

Toucans

Ramphastidae

Toucans, New World barbets are a well defined clade   (Moyle 2004, SACC).

 

Asian Barbets

Megalaimidae

Asian barbets (Megalaimidae) are the basal outgroup to the African barbets and in turn the New World barbets and toucans (Moyle 2004)

 

African barbets

Lybiidae

 
 

Honeyguides

Indicatoridae

Honeyguides (Indicatoridae) and woodpeckers (Picidae) are sister families (Hackett et al. 2008)

 

Woodpeckers

Picidae

 

Passeriformes

Suboscines

   
 

New Zealand Wrens

Acanthisittidae

 
 

Broadbills

Eurylaimidae

The broadbills (family Eurylaimidae) include here both the asities (Philepittidae) of Madagascar and Sapayoa of the New World, and together are the sister group to the pittas (Pittidae) (Irestedt et al. 2006; Moyle 2006a). An alternative classification would be to recognize asities and Sapayoa in their own respective families and two families of broadbills, the Eurylaimidae and Calyptomenidae (TOL)

 

Pittas

Pittidae

The pittas (Pittidae) are sister to broadbills (Eurylaimidae) (Moyle et al 2006)

 

Ovenbirds

Furnariidae

The ovenbird family Furnariidae now includes the phylogenetically embedded woodcreepers, formerly separated as the family Dendrocolaptidae (SACC)

 

Antbirds

Thamnophilidae

 
 

Antthrushes

Formicariidae

 
 

Antpittas

Grallariidae

The antpittas (Grallariidae) are not closely related to the antthrushes (Formicariidae) and thus are treated as a separate family (Chesser 2004, Rice 2005, SACC)

 

Gnateaters

Conopophagidae

 
 

Tapaculos

Rhinocryptidae

 
 

Crescentchests

Melanoparaeiidae

The crescentchests are not related to tapaculos and separated to their own family Melanopareiidae pending resolution of their correct relationships (Chesser 2004, SACC)

 

Tyrant Flycatchers

Tyrannidae

 
 

Sharpbill

move to Tityridae below

Genetic data place Sharpbill in the Tityridae (Ohlson et al. 2008,Tello et al 2009,Harshman 2009)

 

Cotingas

Cotingidae

 
 

Manakins

Pipridae

 
 

Tityras, Becards

Tityridae

The genera placed in the family Tityridae are more closely related to the manakins (Pipridae) than to the cotingas (Cotingidae) (Chesser 2004, Barber and Rice 2007, SACC)

 

Family Uncertain

Incertae Sedis

Includes Philabura and Calyptura; Piprites moved to Tyrannidae (Tello et al 2009)

Passeriformes

Oscines

   
 

Lyrebirds

Menuridae

 
 

Scrubbirds

Atrichornithidae

 
 

Bowerbirds

Ptilonorhynchidae

 
 

Australasian Treecreepers

Climacteridae

 
 

Australasian Wrens

Maluridae

 
 

Honeyeaters

Meliphagidae

 
 

Bristlebirds

Dasyornithidae

 
 

Pardalotes

Pardalotidae

 
 

Australasian Warblers

Acanthizidae

Excludes Mohoua and Finschia moved to Incertae sedis (2.3); (Norman et al 2009); Includes Pachycare (Norman et al 2009)
 

Australasian Babblers

Pomatostomidae

 
 

Logrunners

Orthonychidae

 
 

Satinbirds

Cnemophilidae

 
 

Berrypeckers, longbills

Melanocharitidae

 
 

Painted Berrypeckers

Paramythiidae

Separate the painted berrypeckers (Oreocharis and Paramythia) from the Melanocharitidae to the Paramythiidae  (Barker et al 2004; HBW)

 

Wattled Crows

Callaeidae

 
 

Stitchbird

Notiomystidae

Stitchbird is related to the wattled crows of New Zealand, not to honyeaters (Meliphagidae) (Driskell et al 2007)

 

Whipbirds, Jewel-babblers, quail-thrushes

Psophodidae

Psophodidae includes Androphobus, Psophodes, Ptilorrhoa and Cinclosoma formerly in Eupetidae (now Eupetes only) (Christidis and Boles 2008; see also Cibois et al. 2003, Alstrom et al 2006); resequence to follow Oriolidae

 

Wattle-eyes, Batises

Platysteiridae

 
 

Woodshrikes and allies

Tephrodornithidae

Move Tephrodornis and Philentoma from Incertae Sedis, and Hemipus from Campephagidae to a new shrike family the Tephrodornithidae (Fuchs et al. 2006, 2007)

 

Helmetshrikes

Prionopidae

Separate helmetshrikes (Prionopidae) sister to Vangidae from bushshrikes (Malaconotidae)  (TOL; Beresford et al. 2005; Moyle et al 2006b); if we continue to recognize Vangidae

 

Bushshrikes

Malaconotidae

 
 

Boatbills

Machaerirhynchidae

 
 

Vangas

Vangidae

 
 

Butcherbirds and Allies

Cracticidae

 
 

Bristlehead

Pityriasidae

Bristlehead Pityriasis is related to ioras (Aegithinidae) and perhaps the helmetshrikes (Prionopidae), not to the butcherbirds (Cracticidae) as previously thought (Moyle et al 2006)

 

Woodswallows

Artamidae

 
 

Ioras

Aegithinidae

 
 

Cuckooshrikes

Campephagidae

 
 

Sittellas

Neosittidae

 
 

Whistlers and Allies

Pachycephalidae

Pachycephalidae includes Colluricincla, Pitohui and Oreoica previously separated as Colluricinclidae (Christidis and Boles 2008); remove Hylocitrea, a bombycillid (Spellman et al. 2008)

 

Shrikes

Laniidae

 
 

Vireos, Greenlets

Vireonidae

Vireonidae provisionally includes Erpornis and Pteruthius, new found relatives that may merit family status (Reddy and Cracraft 2007, Reddy 2008)

 

Figbirds, Orioles

Oriolidae

 
 

Drongos

Dicruridae

 
 

Fantails

Rhipiduridae

Includes Silktail (Lamprolia) moved from Monarchidae in 2.3 (Irestedt et al. 2008)
 

Monarchs

Monarchidae

 
 

Crows, Jays

Corvidae

 
 

Australian Mudnesters

Corcoracidae

 
 

Birds-of-paradise

Paradisaeidae

 
 

Australasian Robins

Petroicidae

 
 

Rockfowl

Picathartidae

 
 

Rockjumpers

Chaetopidae

 
 

Rail-babbler

Eupetidae

Restrict Eupetidae to Rail-babbler Eupetes; which is sister to Chaetops and in turn to Picathartes (Jønsson et al. 2007)

 

Waxwings

Bombycillidae

 
 

Silky-flycatchers

Ptilogonatidae

Follow AOU 1998 in recognizing silky flycatchers as a separate family Ptilogonatidae

 

Hypocolius

Hypocoliidae

Separate  Hypocolius as monotypic family Hypocoliidae to be consistent with treatment of other enigmatic waxwing relatives

 

Palmchat

Dulidae

 
 

Oos

Mohoidae

Species of Moho are related to waxwings (Bombycillidae), not to honeyeaters (Meliphagidae) (Fleischer et al. 2008)

 

Hylocitrea

Hylocitreidae

Yellow-flanked Whistler (Hylocitrea) is related to the waxwings, not whistlers (Spellman et al 2008)

 

Fairy Flycatchers

Stenostiridae

Stenostiridae includes Stenostira and Elminia from Incertae Sedis and Culcicapa from Muscicapidae (Barker et al. 2002, 2004; Beresford et al. 2005; Fuchs et al. 2006)

 

Tits, chickadees

Paridae

 
 

Penduline Tits

Remizidae

 
 

Bearded Reedling

Panuridae

Bearded Reedling is not related to parrotbills (Paradoxornis), but is sister to the larks (Alaudidae) and in turn to the rest of the Sylvioidea (Ericson and Johansson 2003; Alström et al. 2006; Fuchs et al. 2006)

 

Nicators

Nicatoridae

Move Nicator species from Incertae Sedis to Nicatoridae, which aligns with Alaudidae and Panuridae to form a basal trichotomy that is sister to the rest of the rest of the sylvioids (Beresford et al 2005, Johansson et al 2008)

 

Larks

Alaudidae

 
 

Bulbuls

Pycnonotidae

 
 

Swallows, martins

Hirundinidae

 

Cettia bush warblers and allies

Cettiidae

See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades

 

Bushtits

Aegithalidae

 
 

Leaf warblers and allies

Phylloscopidae

See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades

 

Reed warblers and allies

Acrocephalidae

See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades

 

Grassbirds and allies

Megaluridae

See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades

 

Black-capped Donacobius

Donacobiidae

Donacobius is related to the Old World warblers, not wrens (Troglodytidae); treated as a monotypic family Donacobiidae (Aleixo and Pacheco 2006; SACC) pending resolution of relationships to Megaluridae vs Bernieridae (Johansson et al 2008)

 

Malagasy warblers

Bernieridae

DNA analyses define this endemic radiation of Malagasy genera previously treated as sylviid warblers and babblers (Cibois et al. 2001; Johansson et al 2008)

 

Cisticolas and allies

Cisticolidae

 
 

Sphenoeacus warbler allies

Incertae Sedis

Includes members of the Sphenoeacus group of Old World warblers separated by Johansson et al (2008)

 

Babblers, Parrotbills

Timaliidae

We defer revision of the babbler complex to a future update that incorporates important papers (Cibois et al 2003, Pasquet et al. 2006, Collar and Robson HBW12, Luo et al 2009; Gelang et al 2009). The Sylvia warblers and white-eyes are embedded in the bab

 

Sylviid Babblers

Sylviidae

We separate Sylvia from other babblers to await better definition of the clade that may include Paradoxornis, Chamea, and (some) Alcippe (Cibois 2003).

 

White-eyes

Zosteropidae

White-eyes are relatives of babblers and possibly subordinated as a subfamily of the Timaliidae when relationships to Yuhina are clarified (Cibois et al. 2003)

 

Sugarbirds and allies

Promeropidae

Expand to include Kakamega (not a babbler), Modulatrix, Arcanator, as well as Promerops (Barker et al 2004, Johanssen et al 2008)

 

Fairy-bluebirds

Irenidae

 
 

Goldcrests, kinglets

Regulidae

 
 

Hyliotas

Hyliotidae

Move hyliotas from the Sylviidae to their own family Hyliotidae, a basal lineage of the Passerida without known modern relatives (Fuchs et al 2006; TOL)

 

Wrens

Troglodytidae

 
 

Gnatcatchers

Polioptilidae

 
 

Nuthatches

Sittidae

 
 

Wallcreeper

Tichodromidae

 
 

Treecreepers

Certhiidae

 
 

Mockingbirds, Thrashers

Mimidae

 
 

Starlings

Sturnidae

Includes Rhabdornis creepers of the Philippines now placed in the Sturnidae (2.3); see Lovette and Rubenstein 2007
 

Oxpeckers

Buphagidae

 
 

Thrushes

Turdidae

 
 

Chats, Old World Flycatchers

Muscicapidae

 
 

Dippers

Cinclidae

 
 

Leafbirds

Chloropseidae

 
 

Flowerpeckers

Dicaeidae

 
 

Sunbirds

Nectariniidae

 
 

Old World Sparrows, Snowfinches

Passeridae

 
 

Weavers, Widowbirds

Ploceidae

 
 

Waxbills, Munias & Allies

Estrildidae

 
 

Indigobirds, Whydahs

Viduidae

 
 

Olive Warbler

Peucedramidae

Peucedramus is sister to Prunellidae (Ericson and Johansson 2003, Jonsson amd Fjeldsa 2006)

 

Accentors

Prunellidae

 
 

Wagtails, Pipits

Motacillidae

 
 

Przevalski's Finch

Urocynchramidae

 
 

Finches

Fringillidae

Fringillidae Includes Hawaiian honeycreeprs as subfamily Drepaninae, following AOU

 

New World Warblers

Parulidae

 
 

Oropendolas, Orioles & Blackbirds

Icteridae

 
 

Bananaquit

Coerebidae

SACC dissolved Coerebidae and placed Coereba along with other genera into Incertae Sedis

 

Buntings, New World Sparrows & Allies

Emberizidae

 
 

Tanagers and Allies

Thraupidae

Boundaries of the Thraupidae versus Emberizidae and Cardinalidae under major revision (NACC, SACC)

 

Longspurs, snow buntings

Calcariidae

Move longspurs (Calcarius) and snow buntings (Plectrophenax) from Emberizidae to their own family Calcariidae (Alström et al 2008)

 

Grosbeaks, Saltators & Allies

Cardinalidae