IOC World Bird List 2.4
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ORDER |
FAMILY |
NOTES ON CLASSIFICATION |
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Tinamidae |
"Early Bird" DNA analyses suggest that flighted tinamous may have evolved from flightless ratites (Hackett et al. 2008) |
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Struthionidae |
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Rheidae |
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Casuariidae |
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Dromaiidae |
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Apterygidae |
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Megapodiidae |
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Cracidae |
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Numididae |
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Odontophoridae |
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Phasianidae |
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Anhimidae |
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Anseranatidae |
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Anatidae |
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Spheniscidae |
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Gaviidae |
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Diomedeidae |
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Procellariidae |
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Hydrobatidae |
The subfamilies Hydrobatinae and Oceanitinae may not be sister taxa and therefore be separate families (Nunn and Stanley 1998; Hackett et al. 2008; Christidis and Boles 2008) |
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Pelecanoididae |
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GrebesGGGrebes |
Podicipedidae |
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Phoenicopteridae |
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Phaethontidae |
The tropicbirds (Phaethontidae) do not belong in the Pelecaniformes and merit treatment as the Order Phaethontiformes (Hackett et al. 2008, Christidis and Boles 2008) |
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Ciconiidae |
The Ciconiiformes now includes only one family; the storks (Ciconiidae); ibises and spoonbills (Threskiornithidae) and herons (Ardeidae) are are moved the Pelecaniformes (Hackett et al 2008) |
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Threskiornithidae |
Ibises and spoonbills (Threskiornithidae) and herons (Ardeidae) are sister taxa in the Order Pelecaniformes (Hackett et al. 2008) |
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Ardeidae |
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Scopidae |
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Balaenicipitidae |
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Pelecanidae |
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| Suliformes | Fregatidae |
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Sulidae |
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Phalacrocoracidae |
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Anhingidae |
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Cathartidae |
SACC elevates the New World Vultures (Cathartidae) to a separate order, the Cathartiformes |
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Sagittariidae |
Secretarybird is best treated as a monotypic family Sagittariidae in accord with the Tree of Life phylogeny (Hackett et al. 2008) |
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Pandionidae |
Ospreys are separated to the family Pandionidae in accord with the Tree of Life phylogeny (Hackett et al. 2008) |
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Accipitridae |
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Falconidae |
The falcons (Falconidae) are not members of the Accipitriformes, but are a separate lineage and Order more closely related to parrots and in turn the Passeriformes (Hackett et al. 2008). Resequence in the future. |
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Otididae |
Move bustards (Otididae) from the Gruiformes to their own Order, the Otidiformes (Hackett et al. 2008) |
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Mesitornithidae |
Move mesites (Mesitornithidae) from the Gruiformes to their own Order, the Mesitornithiformes (Hackett et al. 2008) |
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Cariamidae |
Move seriemas (Cariamidae) from the Gruiformes to their own Order, the Cariamiformes (Hackett et al. 2008) |
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Rhynochetidae |
The Kagu (Rhynochetidae) and Sunbittern (Eurypygidae) are sister families removed from the Gruiformes to a new Order, tentatively named the "Eurypygiformes" (Hackett et al. 2008). |
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Eurypygidae |
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Sarothruridae |
Long recognized to be an artificial assemblage of taxa, the Gruiformes is now restricted to flufftails, finfoots, rails, trumpeters, cranes, and Limpkin (Hackett et al. 2008) |
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Heliornithidae |
The finfoots (Heliornithidae) and flufftails (Sarothruridae) are sister taxa (Hackett et al. 2008) |
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Rallidae |
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Psophiidae |
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Gruidae |
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Aramidae |
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Turnicidae |
The buttonquails (Turnicidae) belong in the Charadriiformes, not the Gruiformes (Hackett et al. 2008) |
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Burhinidae |
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Chionidae |
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Pluvianellidae |
The Magellanic Plover (Pluvianellus) is most closely related to the sheathbills (Chionidae) but treated as a monotypic family by the SACC. |
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Haematopodidae |
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Dromadidae |
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Ibidorhynchidae |
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Recurvirostridae |
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Charadriidae |
The golden plovers (Pluvialis) are a separate lineage that may merit family status (Baker et al 2007, TOL) |
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Pluvianidae |
The Egyptian Plover (Pluvianus) does not belong to the Glareolidae. It is a separate lineage that is the outgroup to plovers, ibisbill, stilts and their allies (Hackett et al. 2008). Therefore separated here to its own family tentatively named "Pluvianidae" |
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Rostratulidae |
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Jacanidae |
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Pedionomidae |
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Thinocoridae |
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Scolopacidae |
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Glareolidae |
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Laridae |
The skimmers and terns are embedded among the gulls in a monophyletic clade; separation of the families Rynchopidae and Sternidae would render the gull family Laridae paraphyletic relative to these other two groups (Baker et al. 2008). |
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Stercorariidae |
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Alcidae |
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Pteroclididae |
The sandgrouse (Pteroclididae) are treated as a separate Order Pteroclidiformes because their relationships to other modern birds remain uncertain. |
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Columbidae |
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Strigopidae |
Parrots (Psittaciformes) are the sister group to the Passeriformes (Hackett et al. 2008); resequence in future. The New Zealand parrots are a basal lineage that is sister to all other parrots; elevate to family status (Christidis & Boles (2008) |
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Cacatuidae |
Cockatoos are treated as a family separate from other parrots (Psittacidae) following Christidis & Boles (2008) |
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Psittacidae |
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Opisthocomidae |
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Musophagidae |
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Cuculidae |
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Tytonidae |
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Strigidae |
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Podargidae |
Defer recognition of the Asian frogmouths (Batrachosomus) vs Australasian frogmouths (Podargus) as separate families (Christidis and Boles 2008) until (basal?) relationships of Rigidipenna inexpectata are resolved (see Tree of Life) |
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Steatornithidae |
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Nyctibiidae |
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Caprimulgidae |
The eared nightjars (Eurostopodus) are the sister taxon to rest of the Caprimulgidae (Barrowclough et al 2006, Larsen et al. 2007), but they are not equivalent in rank to other families in the Caprimulgiformes. |
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Aegothelidae |
The owlet-nightjars (Aegothelidae) are the sister taxon to treeswifts and in turn to the swifts and hummingbirds; hence are included in the Apodiformes (Hackett et al. 2008, Christidis and Boles 2008) |
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Hemiprocnidae |
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Apodidae |
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Trochilidae |
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Coliidae |
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Trogonidae |
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Leptosomidae |
The Cuckoo Roller is the only member of an ancient lineage (not Coraciiformes) with no known relatives among modern birds and thus put into a separate Order, Leptosomiformes (Hackett et al. 2008) |
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Coraciidae |
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Brachypteraciidae |
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Alcedinidae |
The kingfishers (Alcedinidae) comprise three monophyletic clades (Alcedininae, Cerylinae, and Daceloninae) (Moyle 2007); some taxonomists elevate them to family status (Sibley and Monroe 1990, Christidis and Boles 2008) |
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Todidae |
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Momotidae |
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Meropidae |
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Upupidae |
Separation of the hoopoes and hornbills as the Order Bucerotiformes maintains monophyly of the Coraciiformes without submerging the Piciformes and Trogoniformes into the Coraciiformes (Hackett et al. 2008). |
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Phoeniculidae |
The hoopoes (Upupidae) and wood hoopoes (Phoeniculidae) are sister taxa, related in turn to the hornbills (Hackett et al. 2008) |
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Bucerotidae |
The hornbills (Bucerotidae) and ground hornbills (Bucorvidae) are sister taxa, related in turn to the hoopoes and wood hoopoes (Hackett et al. 2008) |
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Bucorvidae |
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Galbulidae |
The jacamars (Galbulidae) and puffbirds (Bucconidae) are well-established sister taxa (SACC, Hackett et al. 2008), with evidence favoring inclusion in the Piciformes, rather than a separate Order Galbuliformes (TOL, Ericson et al. 2006; cf SACC, Fain & Houde 2004) |
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Bucconidae |
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| New World Barbets | Capitonidae | Follow AOU in elevating to Family status as a member of trichotomy with Toucan Barbet and toucans | |
| Toucan Barbet | Semnornithidae | Follow AOU in elevating to Family status as a member of trichotomy with New World barbets and toucans | |
Ramphastidae |
Toucans, New World barbets are a well defined clade (Moyle 2004, SACC). |
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Megalaimidae |
Asian barbets (Megalaimidae) are the basal outgroup to the African barbets and in turn the New World barbets and toucans (Moyle 2004) |
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Lybiidae |
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Indicatoridae |
Honeyguides (Indicatoridae) and woodpeckers (Picidae) are sister families (Hackett et al. 2008) |
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Picidae |
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Acanthisittidae |
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Eurylaimidae |
The broadbills (family Eurylaimidae) include here both the asities (Philepittidae) of Madagascar and Sapayoa of the New World, and together are the sister group to the pittas (Pittidae) (Irestedt et al. 2006; Moyle 2006a). An alternative classification would be to recognize asities and Sapayoa in their own respective families and two families of broadbills, the Eurylaimidae and Calyptomenidae (TOL) |
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Pittidae |
The pittas (Pittidae) are sister to broadbills (Eurylaimidae) (Moyle et al 2006) |
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Furnariidae |
The ovenbird family Furnariidae now includes the phylogenetically embedded woodcreepers, formerly separated as the family Dendrocolaptidae (SACC) |
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Thamnophilidae |
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Formicariidae |
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Grallariidae |
The antpittas (Grallariidae) are not closely related to the antthrushes (Formicariidae) and thus are treated as a separate family (Chesser 2004, Rice 2005, SACC) |
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Conopophagidae |
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Rhinocryptidae |
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Melanoparaeiidae |
The crescentchests are not related to tapaculos and separated to their own family Melanopareiidae pending resolution of their correct relationships (Chesser 2004, SACC) |
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Tyrannidae |
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move to Tityridae below |
Genetic data place Sharpbill in the Tityridae (Ohlson et al. 2008,Tello et al 2009,Harshman 2009) |
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Cotingidae |
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Pipridae |
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Tityridae |
The genera placed in the family Tityridae are more closely related to the manakins (Pipridae) than to the cotingas (Cotingidae) (Chesser 2004, Barber and Rice 2007, SACC) |
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Incertae Sedis |
Includes Philabura and Calyptura; Piprites moved to Tyrannidae (Tello et al 2009) |
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Oscines |
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Menuridae |
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Atrichornithidae |
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Ptilonorhynchidae |
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Climacteridae |
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Maluridae |
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Meliphagidae |
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Dasyornithidae |
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Pardalotidae |
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Acanthizidae |
Excludes Mohoua and Finschia moved to Incertae sedis (2.3); (Norman et al 2009); Includes Pachycare (Norman et al 2009) | ||
Pomatostomidae |
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Orthonychidae |
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Cnemophilidae |
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Melanocharitidae |
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Paramythiidae |
Separate the painted berrypeckers (Oreocharis and Paramythia) from the Melanocharitidae to the Paramythiidae (Barker et al 2004; HBW) |
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Callaeidae |
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Notiomystidae |
Stitchbird is related to the wattled crows of New Zealand, not to honyeaters (Meliphagidae) (Driskell et al 2007) |
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Psophodidae |
Psophodidae includes Androphobus, Psophodes, Ptilorrhoa and Cinclosoma formerly in Eupetidae (now Eupetes only) (Christidis and Boles 2008; see also Cibois et al. 2003, Alstrom et al 2006); resequence to follow Oriolidae |
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Platysteiridae |
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Tephrodornithidae |
Move Tephrodornis and Philentoma from Incertae Sedis, and Hemipus from Campephagidae to a new shrike family the Tephrodornithidae (Fuchs et al. 2006, 2007) |
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Prionopidae |
Separate helmetshrikes (Prionopidae) sister to Vangidae from bushshrikes (Malaconotidae) (TOL; Beresford et al. 2005; Moyle et al 2006b); if we continue to recognize Vangidae |
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Malaconotidae |
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Machaerirhynchidae |
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Vangidae |
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Cracticidae |
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Pityriasidae |
Bristlehead Pityriasis is related to ioras (Aegithinidae) and perhaps the helmetshrikes (Prionopidae), not to the butcherbirds (Cracticidae) as previously thought (Moyle et al 2006) |
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Artamidae |
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Aegithinidae |
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Campephagidae |
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Neosittidae |
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Pachycephalidae |
Pachycephalidae includes Colluricincla, Pitohui and Oreoica previously separated as Colluricinclidae (Christidis and Boles 2008); remove Hylocitrea, a bombycillid (Spellman et al. 2008) |
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Laniidae |
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Vireonidae |
Vireonidae provisionally includes Erpornis and Pteruthius, new found relatives that may merit family status (Reddy and Cracraft 2007, Reddy 2008) |
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Oriolidae |
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Dicruridae |
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Rhipiduridae |
Includes Silktail (Lamprolia) moved from Monarchidae in 2.3 (Irestedt et al. 2008) | ||
Monarchidae |
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Corvidae |
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Corcoracidae |
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Paradisaeidae |
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Petroicidae |
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Picathartidae |
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Chaetopidae |
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Eupetidae |
Restrict Eupetidae to Rail-babbler Eupetes; which is sister to Chaetops and in turn to Picathartes (Jønsson et al. 2007) |
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Bombycillidae |
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Ptilogonatidae |
Follow AOU 1998 in recognizing silky flycatchers as a separate family Ptilogonatidae |
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Hypocoliidae |
Separate Hypocolius as monotypic family Hypocoliidae to be consistent with treatment of other enigmatic waxwing relatives |
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Dulidae |
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Mohoidae |
Species of Moho are related to waxwings (Bombycillidae), not to honeyeaters (Meliphagidae) (Fleischer et al. 2008) |
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Hylocitreidae |
Yellow-flanked Whistler (Hylocitrea) is related to the waxwings, not whistlers (Spellman et al 2008) |
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Stenostiridae |
Stenostiridae includes Stenostira and Elminia from Incertae Sedis and Culcicapa from Muscicapidae (Barker et al. 2002, 2004; Beresford et al. 2005; Fuchs et al. 2006) |
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Paridae |
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Remizidae |
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Panuridae |
Bearded Reedling is not related to parrotbills (Paradoxornis), but is sister to the larks (Alaudidae) and in turn to the rest of the Sylvioidea (Ericson and Johansson 2003; Alström et al. 2006; Fuchs et al. 2006) |
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Nicatoridae |
Move Nicator species from Incertae Sedis to Nicatoridae, which aligns with Alaudidae and Panuridae to form a basal trichotomy that is sister to the rest of the rest of the sylvioids (Beresford et al 2005, Johansson et al 2008) |
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Alaudidae |
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Pycnonotidae |
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Hirundinidae |
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Cettiidae |
See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades |
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Aegithalidae |
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Phylloscopidae |
See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades |
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Acrocephalidae |
See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades |
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Megaluridae |
See Alström et al. (2006) and Johansson et al (2008) for revisions of Old World warbler clades |
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Donacobiidae |
Donacobius is related to the Old World warblers, not wrens (Troglodytidae); treated as a monotypic family Donacobiidae (Aleixo and Pacheco 2006; SACC) pending resolution of relationships to Megaluridae vs Bernieridae (Johansson et al 2008) |
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Bernieridae |
DNA analyses define this endemic radiation of Malagasy genera previously treated as sylviid warblers and babblers (Cibois et al. 2001; Johansson et al 2008) |
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Cisticolidae |
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Incertae Sedis |
Includes members of the Sphenoeacus group of Old World warblers separated by Johansson et al (2008) |
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Timaliidae |
We defer revision of the babbler complex to a future update that incorporates important papers (Cibois et al 2003, Pasquet et al. 2006, Collar and Robson HBW12, Luo et al 2009; Gelang et al 2009). The Sylvia warblers and white-eyes are embedded in the bab |
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Sylviidae |
We separate Sylvia from other babblers to await better definition of the clade that may include Paradoxornis, Chamea, and (some) Alcippe (Cibois 2003). |
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Zosteropidae |
White-eyes are relatives of babblers and possibly subordinated as a subfamily of the Timaliidae when relationships to Yuhina are clarified (Cibois et al. 2003) |
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Promeropidae |
Expand to include Kakamega (not a babbler), Modulatrix, Arcanator, as well as Promerops (Barker et al 2004, Johanssen et al 2008) |
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Irenidae |
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Regulidae |
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Hyliotidae |
Move hyliotas from the Sylviidae to their own family Hyliotidae, a basal lineage of the Passerida without known modern relatives (Fuchs et al 2006; TOL) |
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Troglodytidae |
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Polioptilidae |
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Sittidae |
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Tichodromidae |
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Certhiidae |
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Mimidae |
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Sturnidae |
Includes Rhabdornis creepers of the Philippines now placed in the Sturnidae (2.3); see Lovette and Rubenstein 2007 | ||
Buphagidae |
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Turdidae |
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Muscicapidae |
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Cinclidae |
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Chloropseidae |
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Dicaeidae |
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Nectariniidae |
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Passeridae |
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Ploceidae |
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Estrildidae |
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Viduidae |
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Peucedramidae |
Peucedramus is sister to Prunellidae (Ericson and Johansson 2003, Jonsson amd Fjeldsa 2006) |
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Prunellidae |
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Motacillidae |
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Urocynchramidae |
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Fringillidae |
Fringillidae Includes Hawaiian honeycreeprs as subfamily Drepaninae, following AOU |
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Parulidae |
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Icteridae |
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Coerebidae |
SACC dissolved Coerebidae and placed Coereba along with other genera into Incertae Sedis |
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Emberizidae |
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Thraupidae |
Boundaries of the Thraupidae versus Emberizidae and Cardinalidae under major revision (NACC, SACC) |
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Calcariidae |
Move longspurs (Calcarius) and snow buntings (Plectrophenax) from Emberizidae to their own family Calcariidae (Alström et al 2008) |
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Cardinalidae |
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