"Wisdom begins with putting the right name on a thing"
(Old Chinese Proverb)

Todies, motmots, bee-eaters, hoopoes, wood hoopoes & hornbills

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#,† IOC English name Scientific name Authority Breeding Range Code Comment

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5 Todies Family Todidae

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Todus Brisson, 1760

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Cuban Tody Todus multicolor Gould, 1837 NA : Cuba

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Broad-billed Tody Todus subulatus Gray, GR, 1847 NA : Hispaniola

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Narrow-billed Tody Todus angustirostris Lafresnaye, 1851 NA : Hispaniola

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Jamaican Tody Todus todus (Linnaeus, 1758) NA : Jamaica

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Puerto Rican Tody Todus mexicanus Lesson, 1838 NA : Puerto Rico

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14 Motmots Family Momotidae

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Hylomanes Lichtenstein, MHK, 1839

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Tody Motmot Hylomanes momotula Lichtenstein, MHK, 1839 LA : s Mexico to nw Colombia

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H. m. chiapensis Brodkorb, 1938 Chiapas (s Mexico)

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H. m. momotula Lichtenstein, MHK, 1839 Veracruz (s Mexico) to Honduras

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H. m. obscurus Nelson, 1911 nw Costa Rica to nw Colombia

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Aspatha Sharpe, 1892

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Blue-throated Motmot Aspatha gularis (Lafresnaye, 1840) MA : s Mexico to Honduras

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Momotus Brisson, 1760

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Russet-crowned Motmot Momotus mexicanus Swainson, 1827 MA : nw Mexico to sw Guatemala

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M. m. vanrossemi Moore, RT, 1932 nw Mexico

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M. m. mexicanus Swainson, 1827 nc and c Mexico

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M. m. saturatus Nelson, 1897 sw Mexico

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M. m. castaneiceps Gould, 1855 c Guatemala

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Blue-crowned Motmot Momotus coeruliceps (Gould, 1836) MA : ne Mexico AS Momotus coeruliceps is one of 5 species in Blue-crowned Motmot complex recognized by Stiles 2009, SACC 412

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Blue-diademed Motmot Momotus lessonii Lesson, 1842 MA : s Mexico to w Panama AS Momotus lessonii is one of 5 species in Blue-crowned Motmot complex recognized by Stiles 2009, SACC 412

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M. l. goldmani Nelson, 1900 se Mexico to n Guatemala

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M. l. exiguus Ridgway, 1912 Campeche and Yucatán (s Mexico)

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M. l. lessonii Lesson, 1842 Chiapas (s Mexico) to w Panama

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Whooping Motmot Momotus subrufescens Sclater, PL, 1853 LA : c Panama, n Colombia, n Venezuela; w Ecuador, nw Peru AS Momotus subrufescens is one of 5 species in Blue-crowned Motmot complex recognized by Stiles 2009, SACC 412; it includes "Silver-banded Motmot" M.argenticinctus as a subspecies following Stiles and SACC

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M. s. conexus Thayer & Bangs, 1906 c Panama to nw Colombia

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M. s. spatha Wetmore, 1946 Guajira Pen. (e Colombia)

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M. s. olivaresi Hernández-Camacho & Romero-Zambrano, 1978 nc Colombia

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M. s. subrufescens Sclater, PL, 1853 n Colombia and n Venezuela

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M. s. osgoodi Cory, 1913 w Venezuela

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M. s. argenticinctus Sharpe, 1892 w Ecuador and nw Peru

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Trinidad Motmot Momotus bahamensis (Swainson, 1838) SA : Trinidad and Tobago AS Momotus bahamensis is one of 5 species in Blue-crowned Motmot complex recognized by Stiles 2009, SACC 412

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Amazonian Motmot Momotus momota (Linnaeus, 1766) SA : Amazonia ENG Change English name of Momotus motmota  to Amazonian Motmot with break up of this species complex (Stiles 2009, SACC 412)

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M. m. momota (Linnaeus, 1766) e Venezuela, the Guianas and n Brazil

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M. m. microstephanus Sclater, PL, 1858 se Colombia, e Ecuador and nw Brazil

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M. m. ignobilis von Berlepsch, 1889 e Peru and w Brazil

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M. m. nattereri Sclater, PL, 1858 ne Bolivia

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M. m. simplex Chapman, 1923 w Brazil to wc Brazil (s of the Amazon)

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M. m. cametensis Snethlage, E, 1912 nc Brazil

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M. m. parensis Sharpe, 1892 ne Brazil

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M. m. marcgravianus Pinto & Camargo, 1961 e Brazil

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M. m. pilcomajensis Reichenow, 1919 s Bolivia, s Brazil and nw Argentina

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Andean Motmot Momotus aequatorialis Gould, 1858 SA : Andes from nc Colombia to w Bolivia AS, ENG M. aequatorialis is one of five species in the Momotus momota complex; includes ssp aequatorialis & chlorolaemus (Stiles 2009, SACC); Change English name from Highland Motmot to Andean Motmot following SACC 412

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M. a. aequatorialis Gould, 1858 wc Colombia and e Ecuador

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M. a. chlorolaemus von Berlepsch & Stolzmann, 1902 e Peru and w Bolivia

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Baryphthengus Cabanis & Heine, 1860

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Rufous Motmot Baryphthengus martii (von Spix, 1824) LA : e Honduras to sw Ecuador, w Amazonia

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B. m. semirufus (Sclater, PL, 1853) e Honduras to nw Colombia and w Ecuador

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B. m. martii (von Spix, 1824) w Amazonia

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Rufous-capped Motmot Baryphthengus ruficapillus (Vieillot, 1818) SA : se Brazil, e Paraguay and ne Argentina

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Electron Gistel, 1848

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Keel-billed Motmot Electron carinatum (Du Bus de Gisignies, 1847) MA : s Mexico to n Costa Rica

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Broad-billed Motmot Electron platyrhynchum (Leadbeater, 1829) LA : e Honduras to sw Ecuador, w Amazonia

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E. p. minus (Hartert, 1898) e Honduras to n Colombia

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E. p. platyrhynchum (Leadbeater, 1829) nw Colombia to sw Ecuador

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E. p. colombianum Meyer de Schauensee, 1950 n Colombia

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E. p. pyrrholaemum (von Berlepsch & Stolzmann, 1902) e Colombia, e Ecuador, e Peru and n Bolivia

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E. p. orienticola Oberholser, 1921 w Brazil

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E. p. chlorophrys Miranda-Ribeiro, 1931 c Brazil

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Eumomota Sclater, PL, 1858

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Turquoise-browed Motmot Eumomota superciliosa (Sandbach, 1837) MA : se Mexico, s Mexico to nw Costa Rica

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E. s. bipartita Ridgway, 1912 s Mexico to w Guatemala

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E. s. superciliosa (Sandbach, 1837) se Mexico

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E. s. vanrossemi Griscom, 1929 c Guatemala

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E. s. sylvestris Carriker & Meyer de Schauensee, 1935 e Guatemala

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E. s. apiaster (Lesson, 1842) El Salvador, w Honduras and nw Nicaragua

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E. s. euroaustris Griscom, 1929 n Honduras

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E. s. australis Bangs, 1906 sw Nicaragua and nw Costa Rica

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27 Bee-eaters Family Meropidae

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Nyctyornis Jardine & Selby, 1830

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Red-bearded Bee-eater Nyctyornis amictus (Temminck, 1824) OR : Malay Peninsula, Sumatra, Borneo

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Blue-bearded Bee-eater Nyctyornis athertoni (Jardine & Selby, 1828) OR : widespread

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N. a. athertoni (Jardine & Selby, 1828) India to Indochina and n Malay Pen.

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N. a. brevicaudatus (Koelz, 1939) Hainan I. (off se China)

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Meropogon Bonaparte, 1850

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Purple-bearded Bee-eater Meropogon forsteni Bonaparte, 1850 AU : Sulawesi

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Merops Linnaeus, 1758

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Black-headed Bee-eater Merops breweri (Cassin, 1859) AF : Ivory Coast, Ghana, s Nigeria and n Central African Republic; Gabon to w South Sudan, c DRCongo snf nw Angola

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Blue-headed Bee-eater Merops muelleri (Cassin, 1857) AF : s Cameroon to w Kenya

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Blue-moustached Bee-eater Merops mentalis Cabanis, 1889 AF : Mali and Sierra Leone to w Cameroon and Bioko I AS Blue-moustached Bee-eater is split from Blue-headed Bee-eater (Marks et al 2007, BLI 4)

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Black Bee-eater Merops gularis Shaw, 1798 AF : c, w

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M. g. gularis Shaw, 1798 Sierra Leone to s Nigeria

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M. g. australis (Reichenow, 1885) se Nigeria to w Uganda and n Angola

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Swallow-tailed Bee-eater Merops hirundineus Lichtenstein, AAH, 1793 AF : widespread

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M. h. chrysolaimus Jardine & Selby, 1830 Senegal to Central African Republic

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M. h. heuglini (Neumann, 1906) ne DR Congo, n Uganda, s Sudan and sw Ethiopia

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M. h. furcatus Stanley, 1814 se DR Congo and Tanzania south to Zimbabwe and Mozambique

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M. h. hirundineus Lichtenstein, AAH, 1793 Angola through Namibia and Botswana to n South Africa

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Little Bee-eater Merops pusillus Statius Müller, 1776 AF : widespread

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M. p. pusillus Statius Müller, 1776 Senegal and Gambia to sw Sudan and n DR Congo

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M. p. ocularis (Reichenow, 1900) e DR Congo, n Uganda and c Sudan to the Red Sea coast

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M. p. cyanostictus Cabanis, 1869 e Ethiopia, w Somalia and e Kenya

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M. p. meridionalis (Sharpe, 1892) s and e DR Congo to w Kenya south to c Angola, Zimbabwe and South Africa

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M. p. argutus Clancey, 1967 sw Angola, Botswana and sw Zambia

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Blue-breasted Bee-eater Merops variegatus Vieillot, 1817 AF : ne, c, wc

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M. v. lafresnayii Guérin-Méneville, 1843 Ethiopia and se Sudan PS The highland ssp M. variegatus lafresnayii may be related to and even conspecific with M. oreobates (Turner, 2010), or treated as a separate allospecies (Redman, comm.)

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M. v. loringi (Mearns, 1915) se Nigeria and Cameroon to Uganda and w Kenya

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M. v. variegatus Vieillot, 1817 Gabon to c DR Congo and n Angola

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M. v. bangweoloensis (Grant, CHB, 1915) e Angola to se DR Congo, Zambia and w Tanzania

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Cinnamon-chested Bee-eater Merops oreobates (Sharpe, 1892) AF : s South Sudan to ne Tanzania; w Uganda and e DRCongo to nw Tanzania

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Red-throated Bee-eater Merops bulocki Vieillot, 1817 AF : ec, c, w PL Turner (2010) suggests that Cinnamon-breasted Bee-eater is best treated as a subspecies of Blue-breasted Bee-eater

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M. b. bulocki Vieillot, 1817 Senegal and Gambia to Central African Republic

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M. b. frenatus Hartlaub, 1854 s Sudan and w Ethiopia to ne DR Congo and n Uganda

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White-fronted Bee-eater Merops bullockoides Smith, A, 1834 AF : Gabon to Kenya, south to s Angola, c South Africa and s Mozambique

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Somali Bee-eater Merops revoilii Oustalet, 1882 AF : e Ethiopia and Somalia to e Kenya

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White-throated Bee-eater Merops albicollis Vieillot, 1817 AF : Mauritania and Senegal to e Sudan, Eritrea, Yemen and Somalia

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Böhm's Bee-eater Merops boehmi Reichenow, 1882 AF : w tanzania to Zambia; e Tanzania to c Mozambique

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Green Bee-eater Merops orientalis Latham, 1801 AF, EU, OR : w AF to Arabian Peninsula to Southeast Asia

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M. o. viridissimus Swainson, 1837 Senegal to n and c Ethiopia

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M. o. cleopatra Nicoll, 1910 Nile Valley (Egypt) to n Sudan

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M. o. cyanophrys (Cabanis & Heine, 1860) s Israel, w and s Arabia

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M. o. muscatensis Sharpe, 1886 c and e Arabia

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M. o. beludschicus Neumann, 1910 s Iraq to nw India

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M. o. orientalis Latham, 1801 w India east to Bangladesh

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M. o. ceylonicus Whistler, 1944 Sri Lanka

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M. o. ferrugeiceps Anderson, 1879 ne India to sc China and Indochina

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Blue-cheeked Bee-eater Merops persicus Pallas, 1773 EU : sw, nw AF

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M. p. persicus Pallas, 1773 Egypt and the Middle East to Kazakhstan and w India

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M. p. chrysocercus Cabanis & Heine, 1860 nw Africa

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Olive Bee-eater Merops superciliosus Linnaeus, 1766 AF : Ethiopia to Namibia, Madagascar

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M. s. superciliosus Linnaeus, 1766 e Africa, Madagascar and the Comoro Is.

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M. s. alternans Clancey, 1971 w Angola and nw Namibia

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Blue-tailed Bee-eater Merops philippinus Linnaeus, 1767 OR : widespread

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M. p. javanicus Horsfield, 1821 India and Sri Lanka to s China and Indochina

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M. p. philippinus Linnaeus, 1767 Philippines

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M. p. celebensis Blasius, W, 1885 Sulawesi and Flores (Lesser Sundas)

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M. p. salvadorii Meyer, AB, 1891 New Guinea and New Britain

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Rainbow Bee-eater Merops ornatus Latham, 1801 AU : Australia

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Blue-throated Bee-eater Merops viridis Linnaeus, 1758 OR : s China to Java and Philippines

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M. v. viridis Linnaeus, 1758 s China to Indochina and the Greater Sundas

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M. v. americanus Statius Müller, 1776 Philippines PS See Collar 2011b; await supplements to numerical index

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Chestnut-headed Bee-eater Merops leschenaulti Vieillot, 1817 OR : widespread

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M. l. leschenaulti Vieillot, 1817 India and Sri Lanka to sc China, Indochina and Malay Pen.

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M. l. quinticolor Vieillot, 1817 Sumatra, Java and Bali

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M. l. andamanensis Marien, 1950 Andaman Is.

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European Bee-eater Merops apiaster Linnaeus, 1758 EU, AF : nw Africa, sw Europe to c Asia; South Africa

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Rosy Bee-eater Merops malimbicus Shaw, 1806 AF : s Nigeria; Gabon to c DRCongo

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Northern Carmine Bee-eater Merops nubicus Gmelin, JF, 1788 AF : Mauritania, Senegal and Gambia to Eritrea, Ethiopia and Somalia

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Southern Carmine Bee-eater Merops nubicoides Des Murs & Pucheran, 1846 AF : Angola to w Tanzania, south to Namibia, Botswana and n South Africa

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ORDER BUCEROTIFORMES PHY Separation of the hoopoes and hornbills as the Order Bucerotiformes maintains monophyly of the Coraciiformes without submerging the Piciformes and Trogoniformes into the Coraciiformes (Hackett et al. 2008).

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4 Hoopoes Family Upupidae PHY The hoopoes (Upupidae) and wood hoopoes (Phoeniculidae) are sister taxa related in turn to the hornbills (Hackett et al. 2008)

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Upupa Linnaeus, 1758

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Eurasian Hoopoe Upupa epops Linnaeus, 1758 EU, OR : widespread

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U. e. epops Linnaeus, 1758 nw Africa and Europe e to sc Russia, nw China and nw India.

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U. e. ceylonensis Reichenbach, 1853 c and s India, Sri Lanka

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U. e. longirostris Jerdon, 1862 ne India to s China, Indochina and n Malay Pen.

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U. e. major Brehm, CL, 1855 Egypt

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U. e. senegalensis Swainson, 1837 Senegal and Gambia to Somalia

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U. e. waibeli Reichenow, 1913 Cameroon to nw Kenya and n Uganda

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African Hoopoe Upupa africana Bechstein, 1811 AF : widespread

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Madagascan Hoopoe Upupa marginata Cabanis & Heine, 1860 AF : Madagascar

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St. Helena Hoopoe Upupa antaios Olson, 1975 AO : St. Helena I. EXT Olson, 1975

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9 Wood Hoopoes Family Phoeniculidae PHY The hoopoes (Upupidae) and wood hoopoes (Phoeniculidae) are sister taxa related in turn to the hornbills (Hackett et al. 2008)

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Phoeniculus Jarocki, 1821

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Forest Wood Hoopoe Phoeniculus castaneiceps (Sharpe, 1871) AF : w, c

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P. c. castaneiceps (Sharpe, 1871) Liberia to Nigeria

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P. c. brunneiceps (Sharpe, 1903) Cameroon to DR Congo and Uganda

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White-headed Wood Hoopoe Phoeniculus bollei (Hartlaub, 1858) AF : w, c

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P. b. bollei (Hartlaub, 1858) Liberia to Central African Republic

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P. b. jacksoni (Sharpe, 1890) e DR Congo to s Sudan and c Kenya

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P. b. okuensis Serle, 1949 s Cameroon

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Green Wood Hoopoe Phoeniculus purpureus (Miller, JF, 1784) AF : widespread

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P. p. senegalensis (Vieillot, 1822) s Senegal to s Ghana

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P. p. guineensis (Reichenow, 1902) n Senegal and Gambia to Chad and Central African Republic

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P. p. niloticus (Neumann, 1903) Sudan to w Ethiopia and ne DR Congo

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P. p. marwitzi (Reichenow, 1906) e Uganda and Kenya to e South Africa

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P. p. angolensis (Reichenow, 1902) Angola and w Zambia to ne Namibia and n Botswana

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P. p. purpureus (Miller, JF, 1784) c and sw South Africa

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Black-billed Wood Hoopoe Phoeniculus somaliensis (Ogilvie-Grant, 1901) AF : e

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P. s. abyssinicus (Neumann, 1903) n Ethiopia and Eritrea

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P. s. somaliensis (Ogilvie-Grant, 1901) se Ethiopia, sw Somalia and ne Kenya

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P. s. neglectus (Neumann, 1905) c and sw Ethiopia

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Violet Wood Hoopoe Phoeniculus damarensis (Ogilvie-Grant, 1901) AF : Angola and Namibia

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Grant's Wood Hoopoe Phoeniculus granti (Neumann, 1903) AF : Kenya

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Rhinopomastus Jardine, 1828

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Black Scimitarbill Rhinopomastus aterrimus (Stephens, 1826) AF : e, c, w, sw

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R. a. aterrimus (Stephens, 1826) Senegal and Gambia to w Sudan

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R. a. emini (Neumann, 1905) c Sudan to ne DR Congo and Uganda

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R. a. notatus (Salvin, 1892) Ethiopia

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R. a. anchietae (Barboza du Bocage, 1892) sw DR Congo, Angola and w Zambia

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Common Scimitarbill Rhinopomastus cyanomelas (Vieillot, 1819) AF : s, se

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R. c. schalowi Neumann, 1900 s Somalia to Zambia, e South Africa and Mozambique

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R. c. cyanomelas (Vieillot, 1819) sw Angola, Namibia, Botswana and nc South Africa

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Abyssinian Scimitarbill Rhinopomastus minor (Rüppell, 1845) AF : e

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R. m. minor (Rüppell, 1845) Ethiopia, Somalia and ne Kenya

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R. m. cabanisi (de Filippi, 1853) s Sudan, sw Ethiopia and n Kenya to n Tanzania

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59 Hornbills Family Bucerotidae PHY The hornbills (Bucerotidae) and ground hornbills (Bucorvidae) are sister taxa related in turn to the hoopoes and wood hoopoes (Hackett et al. 2008)

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Anorrhinus Reichenbach, 1849

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Tickell's Brown Hornbill Anorrhinus tickelli (Blyth, 1855) OR : s Burma, sw Thailand ENG Change  English name of Anorrhinus tickelli to Tickell's Brown Hornbill (BLI) with split of A.austeni.

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Austen's Brown Hornbill Anorrhinus austeni Jerdon, 1872 OR : Assam to sw China and Indochina. AS Anorrhinus austeni is split from A.tickelli (Kemp 1995, Rasmussen & Anderton 2005); BLI recognizes

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Bushy-crested Hornbill Anorrhinus galeritus (Temminck, 1831) OR : Malay Peninsula, Sumatra, Borneo

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Tockus Lesson, 1830

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Crowned Hornbill Tockus alboterminatus (Büttikofer, 1889) AF : e, sc, se

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T. a. alboterminatus (Büttikofer, 1889) w Angola

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T. a. geloensis (Neumann, 1905) sw Ethiopia to w Kenya south to n Angola and n Zambia

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T. a. suahelicus (Neumann, 1905) s Somalia and e Kenya to e South Africa

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Bradfield's Hornbill Tockus bradfieldi (Roberts, 1930) AF : s Angola and n Namibia to w Zimbabwe

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African Pied Hornbill Tockus fasciatus (Shaw, 1812) AF : c, w

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T. f. semifasciatus (Hartlaub, 1855) Senegal and Gambia to s Nigeria

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T. f. fasciatus (Shaw, 1812) se Nigeria to n Angola and Uganda

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Hemprich's Hornbill Tockus hemprichii (Ehrenberg, 1833) AF : Eritrea, Ethiopia and n Somalia to Uganda and c Kenya

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Pale-billed Hornbill Tockus pallidirostris (Hartlaub & Finsch, 1870) AF : Angola and s DRCongo to Tanzania and Mozambique

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T. p. pallidirostris (Hartlaub & Finsch, 1870) Angola, s DR Congo and Zambia (except extreme e)

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T. p. neumanni (Reichenow, 1894) e Zambia to s Tanzania and n Mozambique

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African Grey Hornbill Tockus nasutus (Linnaeus, 1766) AF : widespread

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T. n. nasutus (Linnaeus, 1766) Senegal and Gambia to Ethiopia and c Kenya, sw Arabia

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T. n. epirhinus (Sundevall, 1850) s Uganda and s Kenya to n South Africa

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Monteiro's Hornbill Tockus monteiri Hartlaub, 1865 AF : sw Angola to c Namibia

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Northern Red-billed Hornbill Tockus erythrorhynchus (Temminck, 1823) AF : s Mauritania through Somalia to ne Tanzania ENG Change  English name of Tockus erythrorhynchus to Northern Red-billed Hornbill with multiple splits

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Damara Red-billed Hornbill Tockus damarensis (Shelley, 1888) AF : sw Angola, n Namibia, w Botswana. AS Tockus damarensis is split from T.erythrorhynchus (Kemp & Delport 2002)

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Southern Red-billed Hornbill Tockus rufirostris (Sundevall, 1850) AF : Malawi and Zambia to s Angola and Transvaal. AS Tockus rufirostris is split from T.erythrorhynchus ( Kemp & Delport 2002, cf Chittenden 2007)

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Tanzanian Red-billed Hornbill Tockus ruahae Kemp & Delport, 2002 AF : c Tanzania AS Tockus ruahae is a new taxon recognized in the Red-billed Hornbill complex (Kemp & Delport 2002, Tréca & Érard 2000)

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Western Red-billed Hornbill Tockus kempi Tréca & Érard, 2000 AF : Senegal and Gambia to s Mauritania and w Mali. AS Tockus damarensis is a new taxon recognized in the Red-billed Hornbill complex (Kemp & Delport 2002)

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Southern Yellow-billed Hornbill Tockus leucomelas (Lichtenstein, MHK, 1842) AF : s

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T. l. elegans Hartlaub, 1865 sw Angola

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T. l. leucomelas (Lichtenstein, MHK, 1842) Namibia to w Mozambique and n South Africa

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Eastern Yellow-billed Hornbill Tockus flavirostris (Rüppell, 1835) AF : Eritrea, Ethiopia and Somalia to n Tanzania ENG Change English name of Northern Yellow-billed Hornbill to Eastern Yellow-billed Hornbill to be consistent with other world lists

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Von der Decken's Hornbill Tockus deckeni (Cabanis, 1868) AF : Ethiopia and Somalia to c Tanzania

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Jackson's Hornbill Tockus jacksoni (Ogilvie-Grant, 1891) AF : s South Sudan, sw Ethiopia, ne Uganda and w Kenya

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Black Dwarf Hornbill Tockus hartlaubi Gould, 1861 AF : c, w

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T. h. hartlaubi Gould, 1861 Guinea and Sierra Leone to Gabon and Congo

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T. h. granti (Hartert, 1895) DR Congo and ec Congo to w Uganda

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Red-billed Dwarf Hornbill Tockus camurus Cassin, 1857 AF : Sierra Leone to Ghana; s Nigeria to Uganda, Rwanda, c DRCongo and Gabon

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Tropicranus Sclater, WL, 1922

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White-crested Hornbill Tropicranus albocristatus (Cassin, 1848) AF : c, w

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T. a. albocristatus (Cassin, 1848) Guinea to Ivory Coast

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T. a. macrourus (Bonaparte, 1850) Ivory Coast to Benin

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T. a. cassini (Finsch, 1903) Nigeria to n Angola and w Uganda

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Ocyceros Hume, 1873

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Malabar Grey Hornbill Ocyceros griseus (Latham, 1790) OR : sw India

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Sri Lanka Grey Hornbill Ocyceros gingalensis (Shaw, 1812) OR : Sri Lanka

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Indian Grey Hornbill Ocyceros birostris (Scopoli, 1786) OR : India, Nepal

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Anthracoceros Reichenbach, 1849

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Malabar Pied Hornbill Anthracoceros coronatus (Boddaert, 1783) OR : India, Sri Lanka

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Oriental Pied Hornbill Anthracoceros albirostris (Shaw, 1808) OR : widespread

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A. a. albirostris (Shaw, 1808) India to s China, Indochina and n Malay Pen.

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A. a. convexus (Temminck, 1832) s Malay Pen., Greater Sundas, and nearby islands

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Palawan Hornbill Anthracoceros marchei Oustalet, 1885 OR : Philippines

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Black Hornbill Anthracoceros malayanus (Raffles, 1822) OR : Malay Peninsula, Sumatra, Borneo

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Sulu Hornbill Anthracoceros montani (Oustalet, 1880) OR : Sulu Archipelago

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Buceros Linnaeus, 1758

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Great Hornbill Buceros bicornis Linnaeus, 1758 OR : w India, Himalayas to Sumatra

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Rhinoceros Hornbill Buceros rhinoceros Linnaeus, 1758 OR : Malay Peninsula, Greater Sundas

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B. r. borneoensis Schlegel & Müller, S, 1840 Borneo

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B. r. rhinoceros Linnaeus, 1758 s Malay Pen. and Sumatra

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B. r. silvestris Vieillot, 1816 Java

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Rufous Hornbill Buceros hydrocorax Linnaeus, 1766 OR : Philippines

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B. h. hydrocorax Linnaeus, 1766 Luzon and Marinduque (n Philippines)

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B. h. semigaleatus Tweeddale, 1878 c Philippines

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B. h. mindanensis Tweeddale, 1877 s Philippines

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Rhinoplax Gloger, 1841

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Helmeted Hornbill Rhinoplax vigil (Forster, JR, 1781) OR : Malay Peninsula, Sumatra, Borneo

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Penelopides Reichenbach, 1849

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Sulawesi Hornbill Penelopides exarhatus (Temminck, 1823) AU : Sulawesi

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P. e. exarhatus (Temminck, 1823) n Sulawesi

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P. e. sanfordi (Stresemann, 1932) c, s and e Sulawesi and nearby islands

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Visayan Hornbill Penelopides panini (Boddaert, 1783) OR : Philippines ENG Revise English name of Penelopides panini to avoid confusion since all members of the genus have been called "Tarictic" Hornbills; geographical name conforms to other species.

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P. p. ticaensis Hachisuka, 1930 Ticao I. (nc Philippines)

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P. p. panini (Boddaert, 1783) Masbate, Panay, Sicogon, Pan de Azucar, Guimaras and Negros (c Philippines)

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Luzon Hornbill Penelopides manillae (Boddaert, 1783) OR : n Philippines AS Penelopides manillae is split from P.panini (Kemp 1995, Inskipp et al. 1996, cf Dickinson 2003); BLI recognizes

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P. m. manillae (Boddaert, 1783) Luzon, Marinduque, Catanduanes and adjacent islets

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P. m. subniger McGregor, 1910 Polillo and Patnanongan Is.

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Mindoro Hornbill Penelopides mindorensis Steere, 1890 OR : Mindoro, Philippines AS Penelopides mindorensis is split from P.panini (Kemp 1995, Inskipp et al. 1996, cf Dickinson 2003); BLI recognizes

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Mindanao Hornbill Penelopides affinis Tweeddale, 1877 OR : s Philippines AS Penelopides affinis is split from P.panini (Kemp 1995, Inskipp et al. 1996, cf Dickinson 2003); BLI recognizes

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P. a. affinis Tweeddale, 1877 Mindanao, Dinagat and Siargao

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P. a. basilanicus Steere, 1890 Basilan

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Samar Hornbill Penelopides samarensis Steere, 1890 OR : c Philippines AS Penelopides samarensis is split from P.panini (Kemp 1995, cf Inskipp et al. 1996, Dickinson 2003).

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Berenicornis Bonaparte, 1850

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White-crowned Hornbill Berenicornis comatus (Raffles, 1822) OR : Malay Peninsula, Sumatra, Borneo

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Aceros Hodgson, 1844

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Rufous-necked Hornbill Aceros nipalensis (Hodgson, 1829) OR : ne India to w Thailand and nw Vietnam

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Knobbed Hornbill Aceros cassidix (Temminck, 1823) AU : Sulawesi

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Wrinkled Hornbill Aceros corrugatus (Temminck, 1832) OR : Malay Peninsula, Sumatra, Borneo

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A. c. rugosus (Begbie, 1834) Malay Pen. and Sumatra

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A. c. corrugatus (Temminck, 1832) Borneo

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Walden's Hornbill Aceros waldeni (Sharpe, 1877) OR : Philippines

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Writhed Hornbill Aceros leucocephalus (Vieillot, 1816) OR : Philippines

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Rhyticeros Reichenbach, 1849

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Blyth's Hornbill Rhyticeros plicatus (Forster, JR, 1781) AU : Moluccas to Solomon Islands

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R. p. plicatus (Forster, JR, 1781) s Moluccas

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R. p. ruficollis (Vieillot, 1816) n Moluccas, w Papuan islands and w New Guinea

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R. p. jungei Mayr, 1937 New Guinea (except w) and nearby islands

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R. p. dampieri Mayr, 1934 Bismarck Arch.

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R. p. harterti Mayr, 1934 n Solomon Is.

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R. p. mendanae Hartert, 1924 c and s Solomon Is.

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Narcondam Hornbill Rhyticeros narcondami Hume, 1873 OR : Andaman Islands

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Plain-pouched Hornbill Rhyticeros subruficollis (Blyth, 1843) OR : Burma to Malay Peninsula

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Wreathed Hornbill Rhyticeros undulatus (Shaw, 1812) OR : ne India to Borneo and Java

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R. u. undulatus (Shaw, 1812) Bhutan and ne India to Indochina and Malay Pen., Sumatra, Java and Bali

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R. u. aequabilis Sanft, 1960 Borneo

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Sumba Hornbill Rhyticeros everetti Rothschild, 1897 AU : Lesser Sundas

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Bycanistes Cabanis & Heine, 1860

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Piping Hornbill Bycanistes fistulator (Cassin, 1850) AF : w, c

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B. f. fistulator (Cassin, 1850) Senegal and Gambia to w Nigeria

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B. f. sharpii (Elliot, DG, 1873) se Nigeria to n Angola

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B. f. duboisi Sclater, WL, 1922 Central African Republic to c and e DR Congo and w Uganda

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Trumpeter Hornbill Bycanistes bucinator (Temminck, 1824) AF : Angola to Kenya, south to e South Africa

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Brown-cheeked Hornbill Bycanistes cylindricus (Temminck, 1831) AF : Sierra Leone to Ghana

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White-thighed Hornbill Bycanistes albotibialis (Cabanis & Reichenow, 1877) AF : Benin and s Nigeria to w Uganda, c DRCongo and nw Angola

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Black-and-white-casqued Hornbill Bycanistes subcylindricus (Sclater, PL, 1871) AF : c, w

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B. s. subcylindricus (Sclater, PL, 1871) Sierra Leone to w Nigeria

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B. s. subquadratus Cabanis & Schutt, 1880 e Nigeria to w Kenya nw Tanzania and e DR Congo, also n Angola

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Silvery-cheeked Hornbill Bycanistes brevis Friedmann, 1929 AF : South Sudan and Ethiopia to Mozambique

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Ceratogymna Bonaparte, 1854

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Black-casqued Hornbill Ceratogymna atrata (Temminck, 1835) AF : Sierra Leone to Togo; s Nigeria to s South Sudan, c DRCongo and Angola ENG Simplify English name to Black-casqued [Wattled] Hornbill

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Yellow-casqued Hornbill Ceratogymna elata (Temminck, 1831) AF : s Senegal to Cameroon ENG Simplify English name to Yellow-casqued [Wattled] Hornbill

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2 Ground Hornbills Family Bucorvidae PHY The hornbills (Bucerotidae) and ground hornbills (Bucorvidae) are sister taxa related in turn to the hoopoes and wood hoopoes (Hackett et al. 2008)

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Bucorvus Lesson, 1830

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Abyssinian Ground Hornbill Bucorvus abyssinicus (Boddaert, 1783) AF : Senegal, Gambia and Guinea to Ethiopia and Kenya

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Southern Ground Hornbill Bucorvus leadbeateri (Vigors, 1825) AF : Kenya and Tanzania to c Angola, n Botswana and e South Africa